This is a great response, I think this is where I'm confused about the VMX devices. If I deploy the VMX's with Azure public DNS servers for the vnet they're deployed in, they are happy but the clients using the Meraki VPN tunnel cannot reach my internal DNS servers (sitting on-prem behind MX95's). If I change the Azure vnet to look at my internal DNS servers (then reboot the Azure clients), then everything works perfectly -- but once the VMX's are rebooted, they start using those internal DNS servers and then they are unhappy. I think I have a fundamental misunderstanding of how this is supposed to work, but I don't really know how to proceed.
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