The issue never resolved for us so long as we were using the Mail app and had Exchange accounts configured via Meraki. We tried each of the solutions people in here proposed to no effect. Any Exchange accounts that were manually configured on the phones could send email, but not Meraki-pushed accounts. The issue resolved for us when we moved off of the Mail app to the Outlook app for mail. Once Outlook was configured, we were once again able to send images and scanned PDFs (from the Notes app) using the phone's default Share function.
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