Yes they apply to the traffic of AutoVPN too. However the inbound firewall rules list is a bogus list. It isn't actually getting applied. See docs: Note - Inbound Firewall Rules Note that there is currently a section for inbound firewall rules displayed in the Meraki dashboard. However, inbound firewall rules cannot be configured, and this is an error which will be resolved in a future dashboard update. Any rules saved in this field will not be preserved and will have no effect.   Source: https://documentation.meraki.com/MX/Site-to-site_VPN/Site-to-site_VPN_Firewall_Rule_Behavior ... View more